The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.
Gramma was described as "a quiet and constant presence at the zoo, a witness to history, a beloved icon, and an extraordinary ...
Gramma, the San Diego Zoo’s oldest resident and one of the world’s oldest Galapagos tortoises, has died at the estimated age ...
She had “ongoing bone conditions related to advanced age” that recently progressed and the care team “made the compassionate ...
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