Gramma the Galapagos tortoise, oldest resident of San Diego Zoo, dies at about 141

The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has ...

Gramma the Galápagos tortoise, an original San Diego Zoo resident, dies at 141

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after suffering from progressive bone deterioration because of her age.